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Author Topic:   the phylogeographic challenge to creationism
Faith 
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From: Nevada, USA
Joined: 10-06-2001


Message 26 of 298 (263101)
11-25-2005 2:26 PM
Reply to: Message 25 by robinrohan
11-25-2005 10:18 AM


Even if no creationists post replies on this thread, I hope it will be a useful repository for information on the phylogeographic challenge to creationism.
I don't think the creationists see the significance. I'm having a problem myself. What is the relationship between microevolutionary processes maintaining a status quo and "macroevolution"?
I agree Robin. I read through the OP, and it is impressively thorough and thoughtful, but I fail to grasp how it is in any way a challenge to creationism.
I've avoided this thread because I'm afraid it would just bury me in a morass of semantic distinctions, but I guess I'll risk saying this much.

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Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 27 of 298 (263104)
11-25-2005 2:48 PM
Reply to: Message 16 by Parasomnium
11-23-2005 2:39 AM


Very informative, and a great example, both of how micro- and macroevolution are actually just gradations of the same process, and of how to present a decent topic.
How does the OP demonstrate that micro and macro evolution are just gradations of the same process? It is merely a description of the effects of isolating portions of a population which is hardly unknown to creationists. It is what happens in the development of races of human beings too. That is, subgroups of a population take a portion of the gene pool with them, reducing their genetic variability in relation to the parent population, and this develops distinctions in the group from the parent group and from other isolated groups.
This occurs in all the forms of "evolutionary processes." It occurs in natural selection and it occurs in artificial selection (breeding), it occurs for geographic reasons and it occurs for behavioral reasons etc. etc. etc. It occurs wherever a part of a gene pool is isolated reproductively from the larger gene pool, in any way whatever and for any reason whatever, by removing some genetic potentials and bringing new genetic combinations to phenotypic expression that were suppressed in the parent population with its greater genetic variability.
This message has been edited by Faith, 11-25-2005 08:20 PM

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 Message 29 by mick, posted 11-26-2005 4:51 PM Faith has replied

  
Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 33 of 298 (263459)
11-27-2005 3:07 AM
Reply to: Message 32 by mick
11-26-2005 7:08 PM


Re: faith, diversity, f-statistics and sticklebacks
Hi Jar,
You're dead right to pick me (and Faith) up on those points because they rely on a lot of unstated assumptions. It's a very good question. But I think Faith's words are basically okay.
Thanks, Mick, I appreciate very much that you didn't get hung up on my no doubt less than technically exact language, but simply understood what I was saying. I can't always get the right term matched with what I have in mind but I think I can make it reasonably clear if you can continue in this vein.
faith writes:
by removing some genetic potentials
When faith says that, she is assuming that we measure genetic diversity by the number of alleles present in the migratory population compared to the number of alleles present in the ancestral population. If the ancestral population is "very large" and the migrant population is "very small" then it stands to reason that the migrant population, as a subset of the ancestral population, will have fewer alleles and less genetic diversity. It's possible that the migrant population will happen to contain all of the alleles of the ancestral population, but it's unlikely to the extent that the former is smaller than the latter. And it's impossible that new alleles were generated along the way.
This is the "bottleneck" effect.
Thank you. Quite clear.
[AS a side note, the founder effect comes from a different idea. Let's say the ancestral population has three different insulin alleles (A,B and C). Of all the alleles in the ancestral population, 90% are of type A, 4% are of type B and 1% of type C. It is possible that, just by chance, the migrant population has completely different frequencies of each allele. In the migrant population, 30% has allele A, 35% has allele B and 35% has allele C. The migrant population would then have a greater proportion of previously rare alleles, but the alleles wouldn't be "more diverse".]
Again, nicely clear. Appreciate the description. Also nice to learn the terminology "migrant" and "ancestral" for these purposes.
The short of it is that it's not a meaningless question, and Faith is correct to say that colonization of a new habitat by a minority from a large ancestral population will generally involve a reduction of diversity, whether it is measured as the reduction of the number of alleles present in a single population, or a relative reduction of heterozgosity measured across both ancestral and colonizing populations.
Thank you again. It helps to get the specifics laid out as you are doing.
faith writes:
bringing new genetic combinations to phenotypic expression that were suppressed in the parent population
Faith must be talking about dominance here; if heterozygosity if reduced by nonrandom mating then the frequeny of (previously rare) homozygotes increases in the new population, and you get homozygous-recessive phenotypes in the colonizing population that are not expressed in the ancestral population. Right, Faith?
Yes, I often use untechnical language but yes indeed that is what I am thinking of. That's how you could get a blue-eyed migrant population of human beings out of a dark-eyed ancestral population.
Overall a very informative post.
This message has been edited by Faith, 12-01-2005 01:41 AM

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Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 34 of 298 (263460)
11-27-2005 3:45 AM
Reply to: Message 29 by mick
11-26-2005 4:51 PM


Faith writes:
It is merely a description of the effects of isolating portions of a population which is hardly unknown to creationists. It is what happens in the development of races of human beings too. That is, subgroups of a population take a portion of the gene pool with them, reducing their genetic variability in relation to the parent population, and this develops distinctions in the group from the parent group and from other isolated groups.
This occurs in all the forms of "evolutionary processes." It occurs in natural selection and it occurs in artificial selection (breeding), it occurs for geographic reasons and it occurs for behavioral reasons etc. etc. etc. It occurs wherever a part of a gene pool is isolated reproductively from the larger gene pool, in any way whatever and for any reason whatever, by removing some genetic potentials and bringing new genetic combinations to phenotypic expression that were suppressed in the parent population with its greater genetic variability.
Well we agree on all of that.
Wonderful. I'm SO glad that was clear.
Can I just clarify?
Do we agree that, at least in principle, different species of Tamias have different alleles, and different allele frequences, because of phylogeographic processes as you have described them?
Yes. But I have a problem with the term "species." But I know what you mean so I just want to mention it.
Do we agree that these alleles are responsible for the existence of divergent phenotypes in each species (And subpopulation)?
Yes.
Do we agree that some of these phenotypic differences (such as changes in body size, coloration, etc) are diagnostic of the different species?
Don't like that term "species" as I said. How about "subspecies" or "breeds" or "varieties?"
Do we agree that differences in mating preference (if any) might result from such processes?
Could, certainly.
Do we agree then that the diversification of species can be explained with reference to microevolutionary processes?
Problem with term "species." Diversification of varieties/breeds/subspecies certainly may be the result of all these processes.
Finally: can these diagnostic differences between Tamias species be considered "macroevolution" and if not, why not?
They are still Tamias. All species have always been known to vary. Domestic breeders have always produced varieties. The most striking variations are produced by the severest isolation and inbreeding, processes that involve less and less genetic diversity and hence less capacity to vary further without reintroducing the ancestral genes -- and in some cases extreme genetic specialization ("structure?") leads to inability to breed with other varieties of the species, leaving it even more out on a limb as far as future developments of the variety go. So as a principle or rule, overall the production of new phenotypes is accompanied by reduction in genetic diversity which is a trend away from evolutionary potential. I know there are exceptions.
So in other words, the very genetic mechanisms that produce these variations in principle have a built-in limit. At its extreme that limit is reached in actuality by such a "species" as the cheetah whose genetic diversity is almost nil, and whose prospects of survival let alone capacity for further genetic diversity are severely compromised. To my mind this IS the natural barrier to evolution beyond the species. it is built in to the gene pool that it cannot vary beyond a certain point in any direction.
What magnitude of phenotypic change above the origin of species is necessary for you to admit that macroevolution has occurred?
I think there is a huge range of PHENOtypic change that can occur within one species and I don't think I could define it beyond saying that for instance among dogs you'd have to produce one that had none of the dogness of dogs left, and every breed of dog I know of is always characterized by dogness. Same with catness. To some extent I think these qualities can be specified objectively but I'm not sure how far.
I think the evidence would have to be on the genetic level. I would have to see a genetic mechanism that truly and convincingly counteracts the above effect of the reduction of genetic diversity as a result of what is often considered to be "speciation." And not only counteracts but far outstrips it by multiplying genetic diversity to a luxuriant degree. Mutation is usually offered as this mechanism, but it looks to me to be a pretty anemic candidate.
This message has been edited by Faith, 12-01-2005 01:47 AM

This message is a reply to:
 Message 29 by mick, posted 11-26-2005 4:51 PM mick has replied

Replies to this message:
 Message 35 by Modulous, posted 11-27-2005 8:34 AM Faith has replied
 Message 36 by crashfrog, posted 11-27-2005 1:42 PM Faith has replied
 Message 100 by mick, posted 11-28-2005 8:42 PM Faith has replied

  
Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 37 of 298 (263504)
11-27-2005 1:58 PM
Reply to: Message 35 by Modulous
11-27-2005 8:34 AM


Re: Dogness
I don't think I could define it beyond saying that for instance among dogs you'd have to produce one that had none of the dogness of dogs left
That's not large scale evolution though. Nested hierarchy and all that, dogs will always give birth to something with 'dogness' in the same way that mammals will always give birth to something with 'mammalness' and vertebrates always give birth to something with 'vertebrateness'. That is to say that both creationists and evolutionists agree that this process doesn't happen, so it can't be macroevolution.
Well, I was giving the only criterion I could think of for a phenotypic change that would convince me of macroevolution, and if it's impossible it's impossible.
Do we agree that most evolutionary processes (migration, bottleneck, founder effect, natural selection etc etc) do involve a reduction in population accompanied by a reduction in genetic diversity, to one degree or another? So we further agree that it is these processes that eventually produce the new "species" that are observed by biologists? If this is the case, it does seem to me that either these aren't evolutionary processes but devolutionary processes that define the limits of the Kind or Biblical version of a Species -- OR that evolutionists could demonstrate at the extremes of one of these trends a describable loss of the character of the species. Even dogness or catness as many times these animals have been bred out to a genetic limit or near limit which is where any such change according to evolutionary theory should begin to show up it seems to me.
I would have to see a genetic mechanism that truly and convincingly counteracts the above effect of the reduction of genetic diversity as a result of what is often considered to be "speciation." And not only counteracts but far outstrips it by multiplying genetic diversity to a luxuriant degree. Mutation is usually offered as this mechanism, but it looks to me to be a pretty anemic candidate.
Well, in truth, it would have to be mutation coupled with selection, which we know can lead to an increase in information.
But selection if not always then usually leads to a reduction in genetic diversity despite any increase in information. How does this sort out in favor of macroevolution if each minuscule change through mutation only has a recognizable impact on the phenotype in a population with fewer genetic possibilities?
There are other factors too, such as epigenetics, and maybe further others that are yet to be discovered.
You'd have to explain epigenetics to me.

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 Message 35 by Modulous, posted 11-27-2005 8:34 AM Modulous has replied

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 Message 41 by Modulous, posted 11-27-2005 2:28 PM Faith has replied

  
Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 38 of 298 (263506)
11-27-2005 2:15 PM
Reply to: Message 36 by crashfrog
11-27-2005 1:42 PM


At its extreme that limit is reached in actuality by such a "species" as the cheetah whose genetic diversity is almost nil, and whose prospects of survival let alone capacity for further genetic diversity are severely compromised.
Since you've used this example before, you should know that the genetic diversity of cheetah populations has been steadily increasing for about 10,000 years. That is, until recently, as they've been experiencing a marked reduction in population as a result of expanding human encroachment.
Exactly what is it data-wise that is interpreted as a steady increase in genetic diversity for 10,000 years? I believe you told me once, sorry I have trouble keeping those things in mind. I have to comment, however, that if that is the case, poor cheetah. 10,000 years of an increasing chance at survival and its genetic situation is still about as severely limited as it ever gets.
I think there is a huge range of PHENOtypic change that can occur within one species and I don't think I could define it beyond saying that for instance among dogs you'd have to produce one that had none of the dogness of dogs left, and every breed of dog I know of is always characterized by dogness.
I don't understand what "dogness" is. And species Platonism - the concept that you invoke in this paragraph - has been discredited for hundreds of years.
Well, I was answering the question if there is anything on the phenotypic level that I would take as evidence for macroevolution and that's all I could think of -- the loss of the recognizable character of the species. I said there may or may not be some objective way to define dogness or catness, that I wasn't sure how far that could go. It's just that it is very striking that no matter how bizarre and genetically compromised a breed of dogs you can get, and with dogs in particular they can get awfully far out, they all act like dogs. You can get types that look like anything from a small horse to a bear to a rodent to a muffin, but they all act like dogs.
An individual is not in a certain species because it has the "essence" or "nature" of that species; it's in that species because that's the population with which it can mate.
But that neat definition doesn't help me define what I'd regard as evidence for macroevolution. Inability to mate isn't that evidence for me.

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Replies to this message:
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Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 40 of 298 (263509)
11-27-2005 2:26 PM
Reply to: Message 39 by jar
11-27-2005 2:22 PM


Re: easy question
DOGness.
I don't know. I just know it when I see it. They all wag their tails, how about that for a start. But I don't CARE about this, so let's not get off on it. It was the only answer I could think of that could possibly convince me of macroevolution on the phenotypic level. But as I said, I don't think anything on that level would convince me as there is an enormous range possible.
This message has been edited by Faith, 11-27-2005 02:28 PM

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 Message 39 by jar, posted 11-27-2005 2:22 PM jar has replied

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 Message 43 by jar, posted 11-27-2005 2:58 PM Faith has replied

  
Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 42 of 298 (263514)
11-27-2005 2:41 PM
Reply to: Message 41 by Modulous
11-27-2005 2:28 PM


Re: Dogness
Do we agree that most evolutionary processes (migration, bottleneck, founder effect, natural selection etc etc) do involve a reduction in population accompanied by a reduction in genetic diversity, to one degree or another?
Perhaps, and likewise they involve a subsequent increase in genetic diversity but in different directions.
But only through mutation and I'm not at all convinced that confers any truly beneficial effect overall, certainly that it could do so at any rate that would produce macroevolution.
So we further agree that it is these processes that eventually produce the new "species" that are observed by biologists?
I think we can agree there.
If this is the case, it does seem to me that either these aren't evolutionary processes but devolutionary processes that define the limits of the Kind or Biblical version of a Species
That doesn't follow. The seperation of gene pools is followed by a 'filling the niche' scenario, selection is coupled with mutation. Without taking this into account you cannot make the leap to devolution and 'limits of the Kind'. So it must be the OR...
Filling a niche does not affect the genetic picture, it merely expresses it, and selection is not ALWAYS coupled with mutation, and mutation is hardly a convincing mechanism for the kinds of change needed to overcome the effects of the selection processes that fitted the species to the niche, the reduction in genetic diversity that always defines that condition, let alone provide opportunities for further change beyond the niche.
OR that evolutionists could demonstrate at the extremes of one of these trends a describable loss of the character of the species.
Why should this be the case? Why should there be a describable loss of a character? What wouold be a describable loss of the character? We still bare the character of the first vertabrates, we still bare the character of the first lung breathers, we still bare the character of the first mammals, the first primates and the first hominids. How on earth is an evolutionary scientist meant to demonstrate in 150 years what nature could not do to life in billions?
That's OK. In other words there is nothing at all that can be demonstrated phenotypically that would truly demonstrate macroevolution.
Even dogness or catness as many times these animals have been bred out to a genetic limit or near limit which is where any such change according to evolutionary theory should begin to show up it seems to me.
What is dogness? When would a population of dogs bred for so long lose the character of dogs? Surely it would be easier to rid them of what makes them a population of mammals? Nature has been acting on that group for a longer time already.
I'm happy to drop the idea. It is simply the only thing that occurred to me that I would consider proof of macroevolution on the phenotypic level.
But selection if not always then usually leads to a reduction in genetic diversity despite any increase in information.
Not if there is fecundity, one of the things observed by Darwin, most organisms produce more offspring than survive to produce offspring. That is to say, whilst organisms produce lots and lots of offspring, the population level stays about the same.
Fecundity does not increase genetic diversity. You can have enormous numbers of a particular breed but the same reduced genetic potentials. Again, mutation appears to be the only hope for macroevolution.
I think epigenetics is going to get me too far afield so I'll leave it alone for now, but thanks for the link.
This message has been edited by Faith, 11-27-2005 02:41 PM
This message has been edited by Faith, 11-27-2005 02:45 PM

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Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 45 of 298 (263548)
11-27-2005 5:10 PM
Reply to: Message 43 by jar
11-27-2005 2:58 PM


Re: easy question or Reality according to Faith
So a schipperke is not a dog. Do you agree?
I don't know, jar. Haven't yet worked out the definition you know. It's obviously a dog on all the other characteristics one might think of for dogness. Perhaps we need a checklist. But its not having a tail doesn't mean it doesn't retain the behavioral component in some way. Does it wag its butt?

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 Message 43 by jar, posted 11-27-2005 2:58 PM jar has replied

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 Message 47 by halucigenia, posted 11-27-2005 6:33 PM Faith has replied

  
Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 48 of 298 (263576)
11-27-2005 6:52 PM
Reply to: Message 47 by halucigenia
11-27-2005 6:33 PM


Re: A harder easy question
I'm not following you at all, I'm afraid, [AbE or perhaps I just can't entertain your evolutionist assumptions very far] and yes the dogness/catness idea was a throwaway I don't care to pursue.
And I just about never use the term "macroevolution" as I think the whole idea is wrong, only use it when I'm afraid more confusion would result if I didn't.
There is only built-in variation according to the genetic mechanisms within the Kind (a term that is used by creationists to avoid as much as possible the connotations evolutionists attach to the term "species").
To restate it, this variation is limited, I've come to believe, by the observable fact that the more striking the phenotypic variation from the ancestral type, the more limited the genetic potentials and the genetic diversity in the new population, and therefore the less able to produce new types for further selection. And all this adds up to a REDUCTION in evolutionary potential rather than anything that could possibly further it.
The only thing that is ever proposed to counter this trend is mutation, and I have yet to see anything about mutation that suggests it has even the remotest potential to produce enough viable genetic material to support much "micro" evolution against the de-volutionary trends I'm talking about, let alone "macro."
What are called Evolutionary Processes in most intros to population genetics are in fact DEvolutionary Processes, as their trend is ultimately always toward less genetic potential, less genetic diversity, and at the extremes great vulnerability to various diseases and deformities and ultimately extinction.
This message has been edited by Faith, 11-27-2005 07:14 PM

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 Message 49 by crashfrog, posted 11-27-2005 7:20 PM Faith has replied
 Message 50 by halucigenia, posted 11-27-2005 7:55 PM Faith has replied
 Message 53 by DrJones*, posted 11-27-2005 8:21 PM Faith has replied

  
Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 52 of 298 (263601)
11-27-2005 8:14 PM
Reply to: Message 49 by crashfrog
11-27-2005 7:20 PM


Mutations and proteins and so on
The reduction in genetic diversity that accompanies phenotypic variation is OBSERVED. It is KNOWN. The fact that migrated populations vary from the ancestral population via the reduction of genetic possibilities is KNOWN. Ordinary dominance-recessive patterns are OBSERVED to explain a great deal of it.
You'd have to prove that all those mutations have the effects that are much more easily and naturally accounted for by these known mechanisms I am talking about. You'd have to prove they have always occurred at that rate also, and that they do produce beneficial results. In other words, referring to numbers of mutations and the mechanisms of forming proteins does not prove that any of it has anything to do with what Mick originally described on this thread, and he did not appeal to any of those processes either but described it all in terms of population genetics I could easily grasp. Your information simply adds obfuscation whether that is your intention or not.
This message has been edited by Faith, 11-27-2005 08:15 PM

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 Message 49 by crashfrog, posted 11-27-2005 7:20 PM crashfrog has replied

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Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 54 of 298 (263608)
11-27-2005 8:31 PM
Reply to: Message 50 by halucigenia
11-27-2005 7:55 PM


Re: A harder easy question
Sorry that you are not following me, but put simply - if there was a creature that was part way between two kinds would you agree that this showed a gradation between these two kinds?
That helps. I guess I'd agree, if we had arrived at a good definition of the unique features of Catness and Dogness and a convincing combination could be demonstrated. But the two species don't go back to a common ancestor according to evolutionism in any case.
I could not agree more that the whole idea of macroevolution is wrong, it's just that small changes build up to greater ones over time (it's all micro).
The hard thing to understand about your concept of built in and limited variation is what is the mechanism of this limit. Many others have asked what this might be, there is even a whole thread on that one on this forum.
The mechanism is the Evolutionary Processes themselves, the very processes that sort and select. The predictable reduction in genetic diversity is the mechanism.
As others have pointed out selection and bottle necks etc. that are required mechanisms of evolution are reductions in the currently available variations of a large population, however they need variations to work on, and these variations do tend to build up over time and yes, this is due to mutation.
Not at all. The greater number of variations are due to the shuffling of ordinary Mendelian factors such as dominance and recessiveness.
I just don't see why anyone should find it difficult to conceive that over time these small genetic variations cannot build up to allow further selection to operate on them.
It's not hard at all. It's a very easy kind of idea. But genetics really doesn't support it. Variations don't exactly "build up." Whenever there is a variation in the phenotype there is a corresponding reduction in genetic diversity that allowed it to come to expression. They go together hand in hand, the reduction of genetic diversity and the production of a new phenotype. You don't get fancy breeds without the aggressive elimination of lots of genetic potentials, and the same thing happens in nature when you get a new type that is tightly designed to fit a niche and so on. It is only by eliminating other genetic possibilities that you get the new "species" and this being the case variation or "evolution" beyond the given genetic potentials of the original ancestral species is impossible.
I understand that Darwin himself had similar worries about the trend toward less genetic potential that his natural selection theory proposed, so you are in good company there, however that was before genetic variability was understood.
Darwin didn't know anything about genetic potential did he?
Seems to me the more that is known about genetic variability the clearer this pattern is, that the reduction of variability corresponds with phenotypic change, and there has to be a natural limit to this process.
This message has been edited by Faith, 11-27-2005 08:34 PM

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 Message 63 by DBlevins, posted 11-27-2005 9:26 PM Faith has replied
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Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 55 of 298 (263610)
11-27-2005 8:33 PM
Reply to: Message 53 by DrJones*
11-27-2005 8:21 PM


Re: A harder easy question
Two.
Nope, you didn't understand his question at all. He put it better himself.

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 Message 56 by DrJones*, posted 11-27-2005 8:34 PM Faith has replied

  
Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 58 of 298 (263614)
11-27-2005 8:37 PM
Reply to: Message 56 by DrJones*
11-27-2005 8:34 PM


Re: A harder easy question
There are Two. You have no notion of what a Kind is.

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 Message 56 by DrJones*, posted 11-27-2005 8:34 PM DrJones* has replied

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Faith 
Suspended Member (Idle past 1474 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 65 of 298 (263627)
11-27-2005 9:56 PM
Reply to: Message 61 by crashfrog
11-27-2005 8:52 PM


Re: Mutations and proteins and so on
No, it's not. I mean, you're making this up. How can a variation represent a loss of diversity?
Mick understood and agreed. I think I'll stick to discussing it with him.

This message is a reply to:
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