Do we know if the 3 intermediate mutations before the fourth really 'useful' one added any benefits of their own to the organism? It doesn't seem so from the article although it may be that no-one has looked for any yet.
If nothing useful for the intermediates is found, it's not obvious how natural selection can be claimed for them - or at least the story we tell for the evolution of complex structures like the eye is not an exact analogy.
So we're left with asking why something so apparently improbable happens so regularly - 24:96 or 1 roll of the dice in 4. The skeptic would say that on the evidence, something else is happening which we can't yet explain, but the obvious one is we got the odds wrong - ie the dice is loaded somehow. But we haven't yet shown how.
Life, don't talk to me about life - Marvin the Paranoid Android